Leaf structure

A leaf is built in layers: a waxy cuticle, upper epidermis, palisade and spongy mesophyll, lower epidermis with stomata, and veins. Each layer is adapted to its job.

The leaf is the plant's main organ for photosynthesis, gas exchange and transpiration. Its layered structure is adapted for all three.

Parts and functions

PartFunction
CuticleWaxy, waterproof layer that reduces water loss
Upper epidermisTransparent, letting light through to the mesophyll
Palisade mesophyllPacked with chloroplasts; the main site of photosynthesis
Spongy mesophyllAir spaces for gas exchange
StomataPores that control gas exchange and water loss
Guard cellsPairs of cells that open and close each stoma
VeinsXylem and phloem that transport water and food

How structure suits function

The broad, thin blade gives a large surface area for light and a short distance for gases. The palisade layer near the top catches the most light, the spongy layer allows diffusion, and stomata on the lower surface reduce water loss.

  • The broad, flat blade gives a large surface area for capturing light and absorbing carbon dioxide.
  • The blade is thin, so gases have only a short distance to diffuse between the air spaces and the mesophyll cells.
  • Palisade cells are long and column-shaped, tightly packed with chloroplasts just below the upper surface where they receive the most light.
  • Interconnecting air spaces in the spongy mesophyll link to the stomata, letting carbon dioxide diffuse in to the cells and oxygen diffuse out.
  • A branching network of veins carries water to every part of the blade and removes the glucose made, and the guard cells open and close the stomata so the leaf can exchange gases while limiting water loss.

Related processes

The leaf carries out three linked processes. Photosynthesis takes place mainly in the palisade mesophyll, where chloroplasts use light energy to build glucose from carbon dioxide and water.

Gaseous exchange supplies that carbon dioxide and removes the oxygen made: gases move through the stomata and diffuse through the moist air spaces of the spongy mesophyll. Transpiration is the loss of water vapour through the same stomata; it is a side effect of keeping the stomata open for gas exchange, but it also pulls a stream of water up from the roots through the xylem in the veins.

Common labelling errors

Worked exam-style question

Question. The diagram shows a section through a leaf. (a) Name the tissue that is packed with chloroplasts and is the main site of photosynthesis.

(b) Explain two ways the leaf is adapted to absorb light efficiently. (c) Explain how carbon dioxide in the air outside reaches a palisade cell.

(d) State one way the leaf reduces water loss from its surface.

Model answer. (a) The palisade mesophyll. (b) The blade is broad and flat, giving a large surface area to capture light, and the palisade cells are packed with chloroplasts near the upper surface where the light is strongest, while the upper epidermis and cuticle are transparent so light passes straight through to them.

(c) Carbon dioxide diffuses in through the stomata on the lower surface, moves through the air spaces of the spongy mesophyll, and then diffuses down its concentration gradient into the palisade cell, dissolving in the moist cell surface on the way. (d) The waxy cuticle reduces evaporation from the surface (or: most stomata are on the lower surface, and the guard cells can close them to limit water loss).

Source:SRC-DSKP-EN

Frequently asked questions

How is a leaf adapted for photosynthesis?
A leaf is broad and thin to give a large surface area for absorbing light and a short distance for gases to diffuse. Its palisade mesophyll near the upper surface is packed with chloroplasts to trap the most light, the upper epidermis and cuticle are transparent to let light through, the spongy mesophyll has air spaces for carbon dioxide to reach the cells, and veins bring water and carry away the glucose made.
Why are most stomata on the lower surface of a leaf?
The lower surface is cooler and more shaded than the upper surface, so placing most stomata there lets the leaf take in carbon dioxide and release oxygen while losing less water by evaporation. The upper surface, which faces the sun directly, is left almost free of stomata to reduce water loss.
What is the difference between palisade and spongy mesophyll?
Palisade mesophyll lies just below the upper epidermis; its cells are long, column-shaped and tightly packed with chloroplasts, so it is the main site of photosynthesis. Spongy mesophyll lies below it, has fewer chloroplasts and large air spaces between its rounded cells, so its main job is to allow gases to diffuse to and from the photosynthesising cells.

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