Form 4 · Fundamentals of Biology

Metabolism and Enzymes

Metabolism is the sum of all chemical reactions in a cell, building up (anabolism) and breaking down (catabolism). Almost all of these reactions are controlled by enzymes.

This chapter covers what enzymes are, how they work, the factors that change their activity, and their uses in daily life.

Two other factors also change the rate of an enzyme-catalysed reaction: substrate concentration and enzyme concentration. Increasing either one raises the rate because more collisions between enzyme and substrate can occur, but only up to the point where all the available active sites are already occupied at any instant, after which the rate levels off.

Enzymes are named mostly by adding the suffix '-ase' to the name of their substrate or the reaction they catalyse, such as amylase acting on starch or protease acting on protein. Recognising this naming pattern helps you work out what an unfamiliar enzyme does directly from its name in an exam question.

Anabolic reactions include photosynthesis and protein synthesis, both of which build larger molecules and require an input of energy; catabolic reactions include respiration and digestion, both of which break large molecules into smaller ones and release energy that the cell can use.

Digestive enzymes illustrate how optimum pH varies between locations in the body: pepsin in the stomach works best in strongly acidic conditions, while trypsin in the small intestine works best in mildly alkaline conditions, showing that 'optimum pH' is a property of each individual enzyme rather than a single fixed value for all enzymes.

A rate of reaction is calculated from a graph as the gradient of the curve, or more simply as the amount of product formed, or substrate used up, divided by the time taken; SPM questions often ask for this rate to be calculated directly from a results table rather than only described in words.

Content standards in this chapter

  1. 5.1 Metabolism
  2. 5.2 Enzymes
  3. 5.3 Application of Enzymes in Daily Life

Key concepts

Metabolism
Anabolism builds large molecules from small ones (uses energy); catabolism breaks large molecules down (releases energy).
Enzymes as biological catalysts
Enzymes are proteins that speed up reactions without being used up, and are specific to one substrate.
Lock-and-key model
A substrate fits the enzyme's active site like a key in a lock, forming an enzyme-substrate complex.
Effect of temperature
Rate rises to an optimum (around body temperature) then falls sharply as the enzyme denatures at high temperature.
Effect of pH
Each enzyme has an optimum pH; away from it the rate drops, and extreme pH denatures the enzyme.
Applications
Enzymes are used in detergents, food processing, brewing and medicine.
Effect of substrate concentration
As substrate concentration increases, the rate of reaction increases because more substrate molecules collide with the available active sites; beyond a certain point the rate plateaus because every active site is already occupied at any moment.
Effect of enzyme concentration
As enzyme concentration increases, with excess substrate present, the rate of reaction increases because there are more active sites available; the rate eventually levels off once substrate becomes the limiting factor instead.
Naming enzymes
Most enzyme names end in '-ase' and are built from the substrate they act on or the reaction they catalyse, such as amylase for starch, protease for protein and lipase for fat.
Enzymes in daily life
Biological detergents contain protease and lipase to remove protein and fat stains at lower washing temperatures; the food industry uses enzymes such as pectinase to clarify fruit juice and rennin in cheese-making.
Active site and tertiary structure
An enzyme is a globular protein folded into a precise three-dimensional shape; the active site is a small region of this shape whose contours and chemical groups are complementary to one specific substrate.
Immobilised enzymes
Enzymes can be fixed onto or within an inert support, such as alginate beads, so they can be reused repeatedly and easily separated from the product, which is common in industrial applications like producing lactose-free milk.
Calculating rate from data
Rate of reaction can be found by dividing the change in the amount of product or substrate by the time taken, or by finding the gradient of the steepest part of a graph of amount against time.

How this chapter is examined

SPM Biology is assessed over three papers. Paper 1 has 40 objective questions (40 marks) in 1 hour 15 minutes; Paper 2 carries 100 marks across Sections A, B and C in 2 hours 30 minutes; Paper 3 is the practical, testing science process skills. Content from this chapter can appear in any of them, so lessons drill recall for Paper 1 and structured answers for Paper 2.

Common exam angles

  • Interpreting a graph of enzyme activity against temperature or pH.
  • Explaining denaturation in terms of the active site.
  • Designing or evaluating an experiment on enzyme activity (Paper 3).
  • Sketching and explaining a graph of rate of reaction against substrate concentration.
  • Identifying an enzyme from its name and stating the substrate it acts on.
  • Explaining why biological detergents work well at lower washing temperatures.
  • Explaining why different digestive enzymes have different optimum pH values.
  • Describing an industrial or medical use of immobilised or purified enzymes.
  • Distinguishing metabolism from digestion when asked to define the term precisely.

Common mistakes

What students write: Saying enzymes are 'killed' at high temperature.

What earns the mark: Enzymes are not alive; at high temperature they are denatured, their active site changes shape and no longer fits the substrate.

What students write: Writing that a low temperature denatures enzymes.

What earns the mark: Low temperature only slows enzymes down; activity returns when warmed. Only high temperature and extreme pH denature them.

What students write: Treating enzymes as used up in a reaction.

What earns the mark: Enzymes are catalysts and are unchanged after a reaction, so one enzyme molecule can be reused many times.

What students write: Saying an enzyme works on any substrate.

What earns the mark: Each enzyme is specific, its active site fits only one substrate or a group of similar substrates.

What students write: Saying the rate of reaction keeps rising forever as substrate concentration increases.

What earns the mark: The rate plateaus once all the enzyme's active sites are occupied at any given moment, so adding more substrate beyond this point has no further effect.

What students write: Assuming adding more enzyme always speeds up a reaction.

What earns the mark: Once substrate becomes the limiting factor, adding more enzyme no longer increases the rate, because there is not enough substrate for the extra active sites to bind.

What students write: Writing that lipase breaks down protein.

What earns the mark: Lipase breaks down lipids (fats) into fatty acids and glycerol; protease is the enzyme that breaks down protein.

What students write: Describing enzymes only as substances that 'help' reactions.

What earns the mark: Enzymes are biological catalysts: they lower the activation energy needed for a reaction and speed it up without being permanently changed themselves.

What students write: Assuming every enzyme in the body has the same optimum pH.

What earns the mark: Different enzymes have different optimum pH values suited to where they work, such as pepsin in the acidic stomach and trypsin in the mildly alkaline small intestine.

What students write: Describing an enzyme's shape as fixed and rigid at every temperature.

What earns the mark: An enzyme's active site can flex slightly at its optimum temperature, but a large rise in temperature permanently distorts this shape and denatures the enzyme.

What students write: Reading the rate of reaction off a graph as the final value reached.

What earns the mark: The initial rate is usually taken from the steepest part of the graph near the start, since this is when substrate concentration, and therefore rate, is highest.

What students write: Confusing metabolism with digestion, treating the two words as interchangeable.

What earns the mark: Digestion is only one catabolic process that breaks down food; metabolism is the much broader term covering every chemical reaction in the body, both anabolic and catabolic.

Study this chapter

Processes in this chapter

Frequently asked questions

Why does enzyme activity fall at high temperature?
As temperature rises, activity increases up to an optimum. Beyond that, the heat changes the three-dimensional shape of the enzyme's active site, so the substrate no longer fits. The enzyme is denatured and the reaction rate falls sharply, usually permanently.
What is the lock-and-key model?
It describes how an enzyme is specific. The substrate (key) has a shape complementary to the enzyme's active site (lock), so only the correct substrate can bind to form an enzyme-substrate complex, which is why each enzyme catalyses one type of reaction.
Are enzymes used up when they work?
No. Enzymes are catalysts, so they are not used up or permanently changed by the reaction they speed up. A single enzyme molecule can catalyse the same reaction repeatedly, which is why cells need only small amounts.
Why does the rate of an enzyme reaction level off as substrate concentration increases?
At low substrate concentration, increasing it gives more collisions between substrate and the enzyme's active sites, so the rate rises steeply. Once every active site is occupied at a given instant, however, adding more substrate cannot increase the rate any further, since the enzyme itself has become the limiting factor. The graph therefore rises then levels off into a plateau.
How do biological detergents use enzymes?
Biological detergents contain enzymes such as protease and lipase, which break down protein-based stains, like blood or egg, and fat-based stains respectively into smaller, more soluble molecules that rinse away more easily. Because these enzymes work well at body temperature, the detergent is effective at lower washing temperatures than a non-biological one, which can also save energy.
Why do pepsin and trypsin have different optimum pH values?
Pepsin works in the stomach, where hydrochloric acid keeps conditions strongly acidic, so its active site is shaped to function best at a low pH. Trypsin works in the small intestine, where bicarbonate neutralises the acid from the stomach and conditions become mildly alkaline, so its active site instead functions best at a higher pH. Each enzyme's optimum pH reflects the environment it evolved to work in, not a universal rule for all enzymes.
What is an immobilised enzyme and why is it useful?
An immobilised enzyme is fixed onto or trapped within a support material, such as alginate beads, instead of being free in solution. This allows the enzyme to be reused many times and separated easily from the product, which lowers cost and avoids contaminating the product with enzyme; immobilised lactase, for example, is used industrially to produce lactose-free milk.
How do you calculate the rate of an enzyme-catalysed reaction from a graph?
Rate is calculated as the change in the amount of product formed or substrate used divided by the time taken for that change, giving units such as cm³ of gas per minute. On a graph of amount against time, this corresponds to the gradient of the curve; because the curve is usually steepest near the start, the initial rate is normally measured over the first section of the graph, before the substrate becomes limiting.
What is the difference between anabolism and catabolism?
Anabolism refers to reactions that build larger, more complex molecules from smaller ones, such as joining amino acids into a protein during protein synthesis, and this process requires an input of energy. Catabolism refers to reactions that break large molecules down into smaller ones, such as respiration breaking down glucose, and this process releases energy that the cell can then use for anabolic reactions or other cellular work.

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